Basic Facts About Penguins
Penguins are aquatic, flightless birds that are highly
adapted to life in the water. Their distinct tuxedo-like appearance is called
countershading, a form of camouflage that helps keep them safe in the water.
Penguins do have wing-bones, though they are flipper-like and extremely suited
to swimming. Penguins are found almost exclusively in the southern hemisphere,
where they catch their food underwater and raise their young on land.
Diet
Staples: Krill, fish and squid.
General Facts
- penguins closer to the equator eat more fish
- penguins closer to Antarctica eat more squid and krill.
Population
Larger
penguin species are found in colder climates where their large body mass
enables them to cope with the conditions, while smaller penguins inhabit warmer
climes.
The
penguin species with the highest population is the Macaroni penguin with
11,654,000 pairs. The species with the lowest population is the endangered
Galapagos penguin with between 6,000-15,000 individuals.
Range
Penguins
can be found on every continent in the Southern Hemisphere from the tropical
Galapagos Islands (the Galapagos penguin) located near South America to
Antarctica (the emperor penguin).
Behavior.
Penguins
can spend up to 75% of their lives in the water. They do all of their hunting
in the water. Their prey can be found within 60 feet of the surface, so
penguins have no need to swim in deep water. They catch prey in their beaks and
swallow them whole as they swim. Some species only leave the water for molting
and breeding.
The
emperor penguin breeds in the coldest environment of any bird species; air
temperatures may reach -40° (F/C), and wind speeds may reach 89 miles per hour
(144 km/hr)!
Penguins
are social birds. Many species feed, swim and nest in groups. During the
breeding season, some species form large groups, or “rookeries”, that include
thousands of penguins. Each penguin has a distinct call, allowing individuals
to find their mate and their chicks even in large groups.
Reproduction
Mating Season: Varies depending on the species,
though most breed during spring and summer.
Incubation: Varies from 1 month-66 days
depending on the species.
Number of offspring: King and emperor penguins
lay one egg. All other species of penguin lay two eggs.
Threats to Penguins
Penguins
are threatened by climate change. Penguin populations have decreased by nearly
80 percent in some areas, and the majority of scientists agree that rising
temperature due to climate change is the primary culprit.
In
Antarctica, home to the famous emperor penguin, the annual sea ice melting
season has extended by as much as 3 weeks in recent decades. Less ice means
less habitat for penguins and the loss of critical food, such as shrimp-like
krill, which depend on polar ice to reproduce.
The
penguin that is currently most threatened by climate change is the African
penguin. Most African penguins live on islands off the coast of Africa and feed
on a rich supply of anchovies and sardines that follow a nearby current of cold
water. As the oceans heat up, this stream is moving further away from their
island home.
These
penguins will only swim up to 25 miles away from their homes, so if the current
moves much further they will not be able to reach their food source.
In
addition to global warming and natural predation by sharks, orcas, leopard
seals, sea lions and fur seals, other threats to penguins include impacts on
habitat due to oil spills, pesticides, construction, destruction of habitat due
to introduced herbivores, competition with humans for food and illegal egg
harvesting.
Types of Penguins.
Adelie Penguin
Scientific name: Pygoscelis
adeliae
Size: 5.4 kg (m), 4.7 kg (f)
Nest type: in colonies in the open, use stones
to line nest
Favourite food: krill
No
penguin, indeed no other bird, breeds further south than Adelie Penguins.
Adelie Penguins show a number of specialised adaptations to the cold often
encountered by this species that restrict heat loss.
Identification:
A
medium-sized penguin recognised by its white eye-ring. Feathers on the back of
the head are slightly elongated and can be raised to form a small crest.
Light-coloured individuals occur rarely in some colonies. Immature birds up to
14 months of age differ from adults in having a white rather than black chin
and they lack the white ring around their eyes.
Habits:
Adelie
Penguins breed in colonies from a few dozen to many thousands. Within the
colonies, distinct sub-colonies can be recognised. The nests, depressions on
the ground, are lined with small stones, which help to keep the eggs free of
any melt-water from snow. Stones for building nests are often in high demand.
They are highly gregarious on land and at sea. The availability of accessible
ice-free nesting habitats limits the distribution of this species in the high
Antarctic.
Distribution:
Breeds
from October to February on shores around the Antarctic continent, South
Shetland, South Orkney, South Sandwich, and Bouvetøya Islands. At sea Adelie
Penguins are usually found from the edge of the shelf-ice to the northern
extent of the pack-ice.
Migration and Vagrancy:
Adelie
Penguins are migratory and after breeding do not return to their colonies until
the next spring. Little is know about the non-breeding distribution of this
species. There are only a few records of Adelie Penguins during the Antarctic
winter. Recent work using satellite telemetry indicates that Adelie Penguins
from the Ross Sea leave this area in autumn and migrate about 600 km north of
the Antarctic continent. Juveniles are suspected to travel even further north
than adults. Vagrant birds have been recorded as far north as South Georgia,
Falkland Islands, Kerguelen, Macquarie Island, Tasmania, Heard Island, and the
South Island of New Zealand.
Diet:
They
are highly dependent on krill, usually Euphausia superba, although in some
areas E. crystallorophias constitutes the dominant krill species. Fish and
amphipods can be common in certain locations at certain seasons.
African Penguin
Scientific name: Spheniscus
demersus
Size: 3.3 kg (m), 3.0 kg (f)
Nest type: burrow or under bushes/rocks
Favourite food: small fish
This
is the only penguin breeding in Africa and was probably the first penguin
encountered by Europeans. Numbers declined significantly during the Twentieth
Century and their future has been jeopardized recently by major oil spills.
Identification:
The
only penguin occurring regularly in southern Africa. African Penguins, like
Humboldt Penguins, differ from Magellanic Penguins in that they lack a second
dark breast band (although some African Penguins do have an additional breast
band). The area of naked skin reaches all around the eye and is more extensive
than that in Humboldt Penguins. Immature African Penguins have a grey face and
lack the pied pattern of adults. Adult plumage occurs after 14 months.
Habits:
African
Penguins breed in burrows, rock crevices and under shrubs, often forming large
colonies, with some numbering over ten thousand pairs. Breeding is poorly
synchronised. Their loud braying voice led to them also being called Jackass
Penguins. Birds forage close inshore, especially during the breeding season.
Distribution:
As
the name suggests, the African Penguin is endemic to southern Africa with the
largest concentrations along the Benguela Current, which brings nutrient-rich
water to the west coast of South Africa and Namibia.
Migration and Vagrancy:
Non-migratory.
Vagrants have been found as far north as Setta Cama, Gabon, and Inhaca Island,
Mozambique.
Diet:
Mainly
fish (over 80%), in particular anchovy. Cephalopods and crustaceans are taken
as well, but to a much lesser extent.
Chinstrap Penguin
Scientific name
: Pygoscelis antarctica
Size: 5.0 kg (m), 4.8 kg (f)
Nest type: in colonies in the open
Favourite food: krill
This
delicately coloured bird is arguably the most beautiful of penguins. In
contrast to other adult penguins, apart from Royal Penguins, this species has a
white face. This is separated from the white belly by a thin dark line running
under the lower part of the chin - therefore the name.
Identification:
Chinstrap
Penguins are medium-sized penguins, easily recognised by their white face and
the fine black line across the cheeks. The demarcation between the black and
white lies above the eye, isolating the dark eye in the white plumage. The bill
is black. In contrast to most other penguins, juvenile Chinstraps closely
resemble their parents. Until their first moult, juveniles can be recognised by
dark spotting around the eyes and a slightly shorter bill.
Habits:
Usually
breeds on hillside slopes and rocky outcrops in colonies that sometimes can be
enormous. At the South Shetlands, Chinstrap Penguins often breed amongst other
Pygoscelis penguins, though usually on
steeper slopes. Their marine distribution seems to be linked with the limits of
the continental shelf and the confluence of the Antarctic circumpolar and
Weddell Sea currents.
Distribution:
Intermediate
between the Adelie in the south and the Gentoo Penguin in the north. Breeding
colonies are almost exclusively on the Scotia Arc: that is, around the
Antarctic Peninsula on South Georgia, South Orkney, South Shetland, and South
Sandwich Islands (vast numbers on the latter). Small numbers also breed on
Bouvetøya, Peter First, Heard and Balleny Islands.
Migration and Vagrancy:
Chinstrap
Penguins leave their breeding colonies during winter, probably migrating north
of the pack-ice and stay at sea until the next spring. Non-breeders have been
recorded in Adelie Land, Antarctica and stragglers have reached Australia and
Tierra del Fuego.
Diet:
Chinstrap
Penguins feed almost exclusively on krill (Euphasia superba). Other crustaceans
and fish play a minor role.
Emperor Penguin
Scientific name: Aptenodytes
forsteri
Size: 36.7 kg (m), 28.4kg (f)
Nest type: breed on sea ice in winter, egg
carried on feet
Favourite food: fish and squid
The
Emperor Penguin is a bird of extremes in just about every way. It breeds during
the Antarctic winter and exhibits many adaptations to the extreme cold that
these birds experience when breeding.
Identification:
The
Emperor Penguin is bigger than any other living penguin, standing up to 1.1 m
tall. It is distinguished from the smaller King Penguin by its size, more
robust stature, and a broad pale yellow connection between the orange-yellow
ear patches and the pale yellow upper breast. Immature birds resemble adults
but are smaller and have a white rather than black chin. Ear patches are
whitish, becoming increasingly yellow with age.
Habits:
Breeds
during the Antarctic winter from March to December. Eggs and chicks are
balanced on the feet to prevent them from coming into contact with the ice. No
nests are built, which allows the colony to move around and huddle close
together, providing some protection from the cold. The male is solely
responsible for the two-month incubation of the egg during the heart of winter
in almost continuous darkness. If the female has not returned by the time of
hatching, the male is able to feed the chick for a short time with a “milk”
secreted from the oesophagus.
Distribution:
Breeds
during the Antarctic winter in about 30 colonies around the southern parts of
the Antarctic continent, usually on fast ice. Probably depends a lot upon
polynias – areas of open water surrounded by sea ice – during winter.
Migration and Vagrancy:
Little
is known about post-breeding dispersal or migration. Adults stay close to the
permanent ice for most of their lives. Juveniles equipped with satellite
transmitters, however, migrated as far north as the polar front. Vagrants have
turned up on the South Shetland Islands, Tierra del Fuego, the Falklands, South
Sandwich Islands, Kerguelen Island, Heard Island, and New Zealand.
Diet:
Fish,
cephalopods and krill are taken to varying degrees, though cephalopods and fish
probably represent the major components of the diet, especially in terms of
mass.
Erect-crested Penguin
Scientific name: Eudyptes
sclateri
Size: 5.2 kg (m), 5.1 kg (f)
Nest type: in colonies in the open; nest on
rocks with little to no nesting material to line nests
Favourite food: krill and squid
A
little-known rather bizarre bird with a limited breeding distribution in a very
isolated part the world.
Identification:
Similar
to other crested penguins, in particular Snares and Fiordland Penguins. When
dry on land Erect-crested Penguin can be identified by the upright yellow
feather plumes of their crests. Erect-crested Penguins have a distinct gular
pouch, a more parallel bill, and the yellow supercilium attaches higher on the
bill than in Snares and Fiordland Penguins. Identification at sea is extremely
difficult because feather plumes droop down when wet. Immatures have a pale
yellow supercilium without the long plumes and a mottled grey throat. They can
be distinguished from other crested penguins by the lower supercilium, size and
gular pouch.
Habits:
Erect-crested
Penguins breed on rocky slopes bordering the shore. A few pairs build nests but
most lay their eggs onto the bare rock. After a long courtship period two eggs
are laid but the first, much smaller, A-egg is invariably lost, in most cases
on the same day or before the B-egg has been laid.
Distribution:
In
an arc that characterizes the distribution of crested penguins, from the
Antarctic Peninsula and South America through the sub-Antarctic islands in the
Atlantic and Indian Oceans, Erect-crested Penguins form the terminal species in
the east. They are now restricted to the Bounty and Antipodes Islands, with a
few isolated pairs still breeding on the Auckland Islands. All these sites are
south of the subtropical convergence but well north of the polar front. Until
recently there were also some birds breeding on Campbell Island, but they seem
to have disappeared from there now. Abundant sub-fossil material from the
Chatham Islands has also been attributed to this species.
Migration and Vagrancy:
Erect-crested
Penguins do not come to land after their post-breeding moult and their winter
distribution at sea is unknown. Some birds moult regularly on other
sub-Antarctic Islands south of New Zealand and, less commonly, on the South
Island of New Zealand. Vagrants have been recorded from Northland (North Island
of New Zealand), Tasmania, southern Australia, Heard Island and the Falkland
Islands.
Diet:
Diet
has never been studied in this species, but judging from its long foraging
trips, like other crested penguins they probably live mainly on pelagic
crustaceans and fish.
Fiordland Penguin
Scientific name: Eudyptes
pachyrhynchus
Size: 4.1 kg (m), 3.7 kg (f)
Nest type: in forest under vegetation or rocks;
in caves
Favourite food: fish and squid
The
Fiordland Penguin lives in the temperate rainforest of the southwest coast of
the South Island and Stewart Island, New Zealand, where it is endemic.
Identification:
Similar
to Snares Penguin, with a thick yellow stripe running above the eye and ending
in a dropping plume. Distinguished from Snares Penguin by its larger size, a
series of white streaks on the cheeks and the lack of a fleshy margin at the
base of the bill. Immature birds have a mottled white chin, thinner dull yellow
supercilium and probably cannot be safely distinguished from Snares Penguin.
Habits:
Fiordland
Penguins breed under high rainforest canopy, in dense shrub, under boulders and
in caves. The nests are lined with twigs and grass. Colonies usually consist of
loose groups; nests can be several metres apart. All breeding grounds are north
of the subtropical convergence. However, this oceanfront is close to most
breeding sites and is likely to provide most of the food for breeding birds.
The breeding season begins in June during the austral winter. Males fast for 40
to 45 days from arrival until their first foraging trip. The second-laid egg
hatches several days before the first egg. The smaller chick from the latter
typically dies within a few days due to starvation.
Distribution:
Endemic
to New Zealand. Breeds in the cold rainforest of the southwest coast of the
South Island down to Stewart Island.
Migration and Vagrancy:
Migrates
into the Tasman Sea as indicated by at-sea observations and the occurrence of
moulting birds in eastern Australia. Moulting birds regularly occur also on the
Snares Islands. Vagrants have been recorded on the Chathams, Campbell and
Macquarie Islands, and as far as Western Australia in Australia.
Diet:
From
the limited information that is available it appears that the diet can vary
considerably between locations. A study from Codfish Island found that small
pelagic fish larvae contributed over 80% of the food intake by mass, with the
remaining portion made up by squid. On the west coast of the South Island,
however, squid made up over 80% of the diet, crustaceans 13% and fish only 2%.
Both studies, nevertheless, indicate that Fiordland Penguins mainly forage in
pelagic waters undertaking shallow dives.
Galapagos Penguin
Scientific name: Spheniscus
mendiculus
Size: 2.1 kg (m), 1.7 kg (f)
Nest type: burrow or crevices in lava
Favourite food: small fish
The
northernmost of all penguins, Galapagos Penguins breed right on the equator.
Populations fluctuate heavily under the influence of El Niño to a degree that
the future survival of the species is endangered.
Identification:
They
are the smallest of the Spheniscus
penguins. Distinguished by their relatively large bill and narrow white line
around the face.
Habits:
They
breed mainly in caves or crevices of old lava flows and in burrows.
Distribution:
Endemic
to the Galapagos Islands where it breeds on Isabela, Fernandina and possibly
Bartholomew. Distribution seems to be correlated with the Cromwell Current,
which provides cold nutrient-rich surface water.
Migration and Vagrancy:
Galapagos
Penguins remain close to their breeding islands throughout the year. A record
from the Pacific coast of Panama might well have been ship-assisted.
Diet:
The
main prey items taken by Galapagos Penguin are small fish like mullet and
sardines.
Gentoo Penguin
Scientific name: Pygoscelis
papua
Size: 5.6 kg (m), 5.1 kg (f)
Nest type: in colonies in the open
Favourite food: krill and fish
This
is the most northern penguin of this genus and, in many other respects, the odd
one out. In contrast to Chinstrap and Adelie Penguins, some Gentoo Penguins can
be found around their breeding colonies all year round and they forage much
closer inshore than the other two Pygoscelis
species.
Identification:
Gentoo
Penguins are characterised by a white patch around and behind the eye that
joins on the crown. The orange-red lower mandible is also a distinct feature.
Two subspecies are recognised: a larger form in the sub-Antarctic and a
smaller, but otherwise similar subspecies on the Antarctic Peninsula. Juveniles
are very similar to adults, but the white eye-patch is not connected to their
white eye-rings until they moult at an age of 14 months.
Habits:
Colonies
are usually smaller than those of other Pygoscelis penguins and are less
densely packed. In the sub-Antarctic the nests are often found amongst
tussocks, whereas on the Antarctic Peninsula they nest on stony ice-free areas
and beaches.
Distribution:
Mainly
in the sub-Antarctic, but extending to the Antarctic Peninsula. Breeds on
Staten, Falkland, South Georgia, South Sandwich, South Orkney, South Shetland,
the Antarctic Peninsula, Marion, Prince Edward, Crozet, Kerguelen, Heard, and
Macquarie Islands. There is some evidence that the size of colonies depend on
the local width of the continental shelf, i.e. the available inshore foraging
area.
Migration and Vagrancy:
Gentoo
Penguin can be found near their colonies all year round unless ice prevents
access, as it can in the southern parts of their range. Nevertheless, vagrants
have been found as far north as 43°S on the Argentinean coast as well as in
Australia and New Zealand.
Diet:
Dietary
composition varies between season and locations but generally crustaceans, in
particular krill (euphausiids), are the dominant prey in the southern part of
the range, whereas benthic fish are more commonly caught in lower latitudes.
Squid play only a minor role.
Megadyptes
Yellow-eyed
The
genus Megadyptes contains only a single
species, the Yellow-eyed Penguin, which is endemic to southern New Zealand.
Recent phylogenetic studies have shown Megadyptes
to be the sister taxon of the crested penguins, Eudyptes. Yellow-eyed Penguins
share the strong reddish bill and yellow feathers on the head with the crested
penguins, but differ distinctively in many other respects. Generally they rear
two chicks, they forage close inshore and do not migrate. In contrast to most
other penguins, Yellow-eyed Penguins do not breed in dense colonies, but in
single pairs or loose groups with nests out of sight of each other.
King Penguin
Scientific name: Aptenodytes
patagonicus
Size: 16.0 kg (m), 14.3 kg (f)
Nest type: in colonies in the open, have
territories but no nest
Favourite food: fish, some squid
No
other bird has a longer breeding cycle than King Penguins. They take 14 to 16
months to fledge a single chick. During the winter, chicks may be left to fast
for from one to five months (May to September/October). Adults can rear a
maximum of only two chicks every three years.
Identification:
The
second-largest penguin species, similar in appearance to Emperor Penguin, but
their ranges do not usually overlap. Cheeks are dark orange. The belly is white
but the back is paler than other penguins, more of a grey than black. Immatures
are similar to adults, but with duller facial plumage. Ear patches are pale
yellow rather than orange and the throat is grey-white. Reaches adult plumage
after two years.
Habits:
Dense
colonies, which can number several tens of thousand pairs, are located amongst
tussocks, gently sloping beaches, and sometimes can be over a kilometre inland.
No nest is built, but pairs still maintain territories within pecking distance
of each other.
Distribution:
Restricted
to the sub-Antarctic belt, well north of Emperor Penguins. Breeding colonies
are found on Falkland (re-colonised after extermination), South Georgia,
Marion, Prince Edward, Crozet (over half of the world’s population), Kerguelen,
Heard (re-colonised after extermination), and Macquarie Islands. At sea, King
Penguins are usually found in ice-free waters. Telemetry studies have shown
that they forage particularly along the polar front.
Migration and Vagrancy:
Due
to the extended breeding cycle some birds can be found in the colony at any
time of the year. During winter, adults leave their chicks unattended and may
travel extensively before returning. Stragglers have reached the Antarctic
Peninsula, Mawson, Gough Island, South Africa, southern Australia (including
Tasmania), the North and South Islands of New Zealand, as well as New Zealand’s
sub-Antarctic islands.
Diet:
King
Penguins are specialised on pelagic fish, in particular laternfish of the
species Electrona carlsbergi , Kreffichthys anderssoni and Protomyctophum
tenisoni, which can make up over 99 % of the diet. Cephalopods play a minor
role and, to an even lesser extent, so do crustaceans.
Little Penguin
Scientific name: Eudyptula
minor
Size: 1.2 kg (m), 1.0 kg (f)
Nest type: burrow, cave or under bushes
Favourite food: small fish
The
world’s smallest penguin (also known as Little Blue, Blue and Fairy Penguin).
Identification:
The
Little Penguin closely resembles juveniles of the genus Spheniscus, but their
ranges do not overlap. Upper parts are pale blue to a dark grey-blue depending
upon age, season and subspecies. The transition from the dark upper parts to
the white plumage of the lower body is not as well defined as in other
penguins, going through shades of grey and brown, especially in the face.
Habits:
In
contrast to the other species, Little Penguins are nocturnal. That means they
generally do not enter the shore before dusk and leave it before dawn. They
forage during the day and often will sleep beside the nest at night after they
have fed chicks. This species nests in burrows, under trees, in rock crevices,
and sometimes in caves. Usually nests are clustered to form colonies, but
single breeding pairs are not uncommon. At sea Little Penguins are often found
alone or in small groups of up to ten birds, but sometimes these groups can be
much larger. Although foraging trip durations can be highly variable, Little
Penguins tend to stay close to the coast.
Distribution:
Little
Penguins are widely distributed in Australia (from Western Australia along the
southern coast of Australia up to New South Wales) and in New Zealand (from
Northland to Stewart Island and the Chatham Islands). The White-flippered
Penguin (E. m. albosignata) is an
endangered subspecies, restricted to Banks Peninsula and Motonau Island (South
Island, New Zealand) that has often been treated as a full species. Geographic
variation of size, extent of white on the tail and flipper, and colour tone of
the back is considerable. Six subspecies have been described: novaehollandia in Australia, iredaei in northern New Zealand, variabilis from Cook Strait, New Zealand, albosignata on Banks Peninsula, minor in the lower part of the South Island,
New Zealand, and chathamensis from the
Chatham Islands.
Migration and Vagrancy:
Juveniles
disperse widely after fledging. Adults sometimes undertake long trips at sea
during the non-breeding season, but return regularly to the colony throughout
the year.
Breeding Season and Moult:
The
breeding season is highly variable from place to place and in some areas from
year to year. It usually begins in August/September. The first chicks fledge in
November/December, but in many areas there are second or, sometimes even, third
clutches laid, which can extend the breeding season up to May. Little Penguins
moult once at the end of the breeding season. At Phillip Island, Australia,
this occurs mainly between mid-February and mid-March. Moulting is more
synchronised than laying.
Diet:
Mainly
fish, especially sardines and anchovies, but also cephalopods and to a very
small degree crustaceans.
- Little Penguin chick near fledging. Photo: Lloyd Davis Photography
Macaroni Penguin
Scientific name: Eudyptes
chrysolophus
Size: 5.2 kg (m), 5.3kg (f)
Nest type: in colonies in the open
Favourite food: krill
This
is probably the most abundant of all penguins in terms of total numbers. Brood
reduction is taken to an extreme by this species. The Royal Penguin (Eudyptes chrysolophus schlegeli), a
white-faced variant, is treated as a subspecies of Macaroni Penguin here, but
others accord the Royal penguin full species status.
Identification:
In
contrast to the other crested Penguins, this species has orange, not yellow,
feather plumes. They originate from a supercilium that meets at the front, i.e.
higher up the head than in other species. Macaroni Penguins are also slightly
larger than the other crested penguins. Most Macaroni Penguins breeding on
Macquarie Island (south of Australia) have a white face and are referred to as
Royal Penguins: they are sometimes given full species status (Eudyptes schlegeli), although the biological
basis for doing so is very doubtful. Immatures are similar to adults but lack
the long feather crest. Instead only a short orange-yellow supercilium is
present.
Habits:
Breeds
on rocky slopes, beaches and amongst tussocks. Most birds build a small nest
from pebbles and by scraping out some mud or sand, but many pairs are content
with laying their two eggs on bare rock. Egg-size dimorphism of Macaromi
Penguins (and Erect-crested Penguins) is amongst the largest known for any
bird. The first-laid A-egg is about 61-64% smaller than the B-egg. In most
cases the A-egg is lost before or on the day the B-egg is laid, and it almost
never survives to fledging even though the embryo is viable. Satellite
telemetry studies indicate that Macaroni Penguins forage mainly along the polar
front regularly travelling up to 400 km to reach a feeding site.
Distribution:
The
distribution of Macaroni Penguin extends from the sub-Antarctic to the
Antarctic Peninsula, but overall they are found further south than the rest of
the crested penguins. The range overlaps with that of the southern form of the
Rockhopper Penguin. Breeding colonies are found on the Antarctic Peninsula,
islands around Cape Horn, Falklands, South Georgia, South Sandwich, South
Orkney, South Shetland, Bouvetøya, Prince Edward, Marion, Crozet, Kerguelen,
Heard Island and Macquarie Island.
Migration and Vagrancy:
Macaroni
Penguins are migratory and found only exceptionally near land during the
non-breeding season. Vagrant dark-faced birds are known from South Africa,
Antarctica, Campbell Island, and The Snares. Royal Penguins have been recorded,
possibly breeding, on Heard, Kerguelen, Crozet Island, and Marion Island
amongst dark-faced Macaroni Penguins and stragglers have been observed as far
north as North Island, New Zealand.
Diet:
The
diet is mainly composed of crustaceans. On Macquarie, euphausiids make up only
half the diet by weight, with fish constituting the other half.
Magellanic Penguin
Scientific name: Spheniscus
magellanicus
Size: 4.9 kg (m), 4.6 kg (f)
Nest type: burrow or under bushes
Favourite food: small fish
Similar
to African Penguin (to which it is very closely related), but breeding on the
opposite side of the Atlantic. It is also the only migratory, offshore-foraging
species in this genus.
Identification:
It
is the only Spheniscus penguin found
over most of its range, but overlaps the distribution of Humboldt Penguins
around Puerto Montt, Chile. Humboldt Penguins lack the second dark breast band
found in Magellanic Penguin and have more extensive areas of bare facial skin.
However, as both of these characters are subject to individual variation and
hybrids do occur, not every bird might be identifiable. Some immature birds
undergo partial head moult during winter and gain the pied head pattern of
adults.
Habits:
Breeds
in burrows where digging is possible, otherwise on the surface or under bushes.
Colonies form in a variety of habitats from low forests to grassland to bare
rocks, often on islands or headlands. Some colonies on the Argentinean side
number several hundreds of thousands of pairs.
Distribution:
Breeds
around the southern tip of South America from 40°S in Argentina to 37°S in
Chile, as well as on the Falkland Islands. The largest colonies are found on
the Atlantic side of South America.
Migration and Vagrancy:
Magellanic
Penguins are migratory, some birds moving as far north as Peru and Brazil in
winter. Vagrants have been recorded in South Georgia, on the Antarctic
Peninsula, Australia, and New Zealand.
Diet:
Fish,
mainly anchovies and sardines, are supplemented by cephalopods. Considerable
variation between sites and years has been recorded, however, and one study
found a small crustacean to be the main prey item.
Rockhopper Penguin
Scientific name: Eudyptes
chrysocome
Size: 2.5 kg (m), 2.4 kg (f)
Nest type: in colonies in the open; sometimes
in association with other species
Favourite food: krill, fish and squid
Breeding
in sometimes-large colonies throughout the sub-Antarctic. Smaller than its
congeners, but no less aggressive. There is some evidence that the Northern
Rockhopper or Moseley’s Penguin is deserving of separate species status.
Whatever, the Northern Rockhopper and Southern Rockhopper are clearly closely
related and much of what applies to one probably holds for the other, but
actual data are still scarce.
Identification:
Rockhoppers
are distinguished from other crested penguins by their smaller size and by
having only a thin yellow superscilium. The feather plumes are yellow, not
orange as in Macaroni Penguin, and thinner than in the remaining Eudyptes
species. The red eye is distinctive. Southern Rockhopper Penguins differ from
their Northern counterparts in having a narrower supercilium and shorter
plumes, which reach just over the black throat. Their vocalisations are also
different. The Southern Rockhopper actually comprises two subspecies that have
been described and can be identified in the field: the nominate form from South
America and the Falkland Islands and the eastern subspecies filholi from the
New Zealand sub-Antarctic islands. The eastern form mainly differs from the
nominate subspecies in having a pink line of fleshy skin along the lower
mandible which is black in the nominate subspecies. Immature birds have only a
narrow supercilium and a pale mottled grey chin. Identification of juveniles is
difficult. Shape of the supercilium, bill shape, body size and underwing
pattern can aid identification. Separation of juvenile Southern and Northern
Rockhopper Penguins in the field is probably impossible.
Habits:
Breeding
colonies are located on rocky slopes and amongst tussocks, sometimes in small
caves and amongst crevices. A small nest is build from tussock, peat and
pebbles. Nevertheless, most of the first-laid eggs (A-eggs) are lost during
incubation. The few chicks that hatch from A-eggs almost invariably die during
the first few days of brooding.
Distribution:
The
northern form of the Rockhopper Penguin breeds in cool temperate climates,
generally north of the subtropical convergence, with breeding occurring on
Tristan da Cunha and Gough Island in the Atlantic Ocean and St. Paul and
Amsterdam Islands in the Indian Ocean. The breeding season starts three months
earlier (July) than in the southern form. The latter is restricted to the
northern sub-Antarctic and has a circumpolar distribution. Breeding colonies
are around the Cape Horn area, Falklands, Prince Edward, Marion, Crozet,
Kerguelen, Heard, Macquarie, Campbell, Auckland and Antipodes Islands. Campbell
Island used to be the eastern stronghold of the species, but the population
there has plummeted recently.
Migration and Vagrancy:
The
non-breeding pelagic range is poorly known. Moulting birds especially have been
found in South Africa, Australia and New Zealand. The western subspecies
(nominate form) has been recorded as far as the Snares Islands during moult.
Vagrants of the Northern Rockhopper have been recorded on the Chatham Islands.
Diet:
Crustaceans,
in particular euphausids, make up the bulk of food items consumed during most
studies of this species. Fish and cephalopods play a minor role, though one
study found 53% cephalopods (by weight). Over 90% of the diet (by mass) of
Northern Rockhopper Penguins breeding on Gough Island consisted of crustaceans
(mainly euphausids). The remaining 10% was made up of fish and, to a very small
extent, squid.
Royal Penguin
Scientific name: Eudyptes
schlegeli or Eudyptes chrysolophus
schlegeli
Size: 5.2 kg (m), 5.3kg (f)
Nest type: in colonies in the open
Favourite food: krill
The
Royal Penguin is treated as a separate penguin species (Eudyptes schlegeli) by some. However, other authorities
highlight evidence from DNA, morphology and behaviour that points to Royal
Penguins being a white-faced variant (Eudyptes
chrysolophus schlegeli) of Macaroni Penguins.
Identification:
In
contrast to the other crested penguins, Royal (and Macaroni) Penguins have
orange, not yellow, feather plumes. They originate from a supercilium that
meets at the front; i.e. higher up the head than in other species. These
penguins are also slightly larger than the other crested penguins. Royal
Penguins breed on Macquarie Island and most have a white face. Immatures are
similar to adults but lack the long feather crest. Instead only a short
orange-yellow supercilium is present.
Habits:
Breeds
on rocky slopes, beaches and amongst tussocks. Most birds build a small nest
from pebbles and by scraping out some mud or sand, but many pairs are content
with laying their two eggs on bare rock. Satellite telemetry studies indicate
that Royal penguins forage mainly along the Polar Front regularly travelling up
to 400 km to reach a feeding site.
Distribution:
Breeds
on Macquarie Island.
Migration and Vagrancy:
Royal
penguins have been recorded, possibly breeding, on Heard, Kerguelen, Crozet
Island, and Marion Island amongst dark-faced Macaroni penguins and stragglers
have been observed as far north as North Island, New Zealand.
Diet:
The
diet is mainly composed of crustaceans. On Macquarie, euphausiids make up only
half the diet by weight, with fish constituting the other half.
- Royal Penguins, Macquarie Island. Photo: Lloyd Davis Photography
Snares Penguin
Scientific name: Eudyptes
robustus
Size: 3.3 kg (m), 2.8 kg (f)
Nest type: in colonies in the open or under
forest canopy
Favourite food: krill, squid and fish
Similar
in many respects to Fiordland Penguin but endemic to the Snares Islands, which
are about 100 km south of the nearest Fiordland Penguin breeding sites. With
its breeding range confined to the just over 300 ha of the Snares group, it has
the most restricted distribution of all penguins.
Identification:
Similar
to Fiordland Penguins (see above for differences). Differs from Erect-crested
Penguins in having drooping feather plumes on the crest, the yellow facial
stripe reaches further up the bill, and they have a more conical bill. The
underwing pattern is highly variable and of little use for identification in
the field.
Habits:
Breeds
under the canopy of Olearia forests, as well as on coastal rock. The forest
often dies due to penguin guano, creating clearings. The birds build a cup nest
from peat, wood and pebbles and nest in dense colonies. Similar to Fiordland
Penguin, both chicks usually hatch, but only the first-hatched chick (from the
B-egg) typically survives beyond the first few days.
Distribution:
Endemic
to the Snares Islands south of New Zealand. Little is known about the
non-breeding distribution.
Migration and Vagrancy:
Thought
to move westwards into the Tasman Sea. Vagrants have been recorded on Macquarie
Island and on the Falklands.
Diet:
The
few studies that have been undertaken indicate that this species lives mainly
on euphausiids, but also takes cephalopods and a few fish.
Yellow-eyed Penguin
Scientific name: Megadyptes
antipodes
Size: 5.7 kg (m), 5.4kg (f)
Nest type: under dense vegetation
Favourite food: fish and squid
The
Yellow-eyed Penguin is often referred to as the rarest penguin in the world,
although, unfortunately, there are others that could lay claim to that crown
too: especially the Galapagos and Fiordland Penguins.
Identification:
Adults
are unmistakable with their yellow eyes and yellow eye-stripes that join on the
back of the head. Moulting birds and birds at sea can be confused with crested
penguins. Immature birds are similar to adults but have a pale yellow chin and
a less vivid yellow eye-stripe.
Habits:
They
nest in dense vegetation in dunes and coastal forest, with nests typically
being isolated from each other. At sea, Yellow-eyed Penguins forage in pairs or
alone.
Distribution:
Endemic
to New Zealand, Yellow-eyed Penguins breed on the east and south coast of the
South Island, on and around Stewart Island, the Auckland Islands, and Campbell
Islands.
Migration and Vagrancy:
Resident.
Adults can be found near the breeding colonies throughout the year. Juveniles
disperse as far north as East Cape, North Island, but none have been recorded
in Australia.
Breeding Season and Moult:
Yellow-eyed
penguins have a very long chick-rearing period (100 days). Consequently,
breeding takes from September to February. Moult occurs at the end of the
breeding period.
Diet:
Yellow-eyed
Penguins feed mainly on fish. Both, pelagic and demersal species are taken.
Cephalopods are taken to a lesser extent, but may be more important for
immatures and for adults in years when available fish stocks are limited.